:root{
  --bg:#f6f4ef; --paper:#fffefa; --ink:#202525; --muted:#6d7472; --accent:#315f62;
  --line:#d9d7cf; --soft:#eceee9; --card1:#b8c8c0; --card2:#c7bcae; --card3:#aabdc2; --card4:#c0b5c1;
  --max:1440px; --serif: Georgia, 'Times New Roman', serif; --sans: Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, 'Segoe UI', sans-serif;
}
*{box-sizing:border-box} html{scroll-behavior:smooth} body{margin:0;background:var(--bg);color:var(--ink);font-family:var(--sans);line-height:1.55} img,video{max-width:100%;display:block} a{color:inherit;text-decoration:none} a:hover{color:var(--accent)}
.container{width:min(calc(100% - 48px),var(--max));margin:0 auto}.narrow{width:min(calc(100% - 48px),1100px);margin:0 auto}
.site-header{position:sticky;top:0;z-index:50;background:rgba(255,254,250,.96);backdrop-filter:blur(14px);border-bottom:1px solid var(--line)}
.nav{height:78px;display:flex;align-items:center;justify-content:space-between;gap:24px}.brand{font-size:.95rem;font-weight:800;letter-spacing:.07em}.nav-links{display:flex;align-items:center;gap:27px;font-size:.92rem;color:var(--muted)}.nav-links a.active{color:var(--ink);font-weight:700}.nav-erc{display:inline-flex!important;align-items:center;gap:6px}.nav-erc img{width:25px;height:25px;object-fit:contain}.menu-btn{display:none;border:0;background:transparent;font-size:1.4rem}
.hero{position:relative;height:min(78vh,780px);min-height:590px;background:#090909;overflow:hidden}.hero-slide{position:absolute;inset:0;opacity:0;transition:opacity 1.25s ease}.hero-slide.active{opacity:1}

/* Each hero visual has its own composition. Only slide 1 is full-bleed. */
.hero-slide:nth-child(1) video{width:100%;height:100%;object-fit:cover}

/* DREADD histology: deliberately compact and anchored at right, as in the approved mock-up. */
.hero-slide:nth-child(2) img{position:absolute;right:1.5%;top:4%;width:42%;height:84%;object-fit:contain;object-position:center right}

/* Cortical-stimulation brain: show the complete brain rather than cropping the superior/inferior edges. */
.hero-slide:nth-child(3) img{position:absolute;left:auto;right:3%;top:1.5%;width:52%;height:87%;object-fit:contain;object-position:center right}

/* DigiTrack: retain both low-resolution heat maps at a modest size, clear of the text block. */
.hero-slide:nth-child(4) img{position:absolute;right:1.5%;top:17%;width:55%;height:66%;object-fit:contain;object-position:center right}

.hero::after{content:'';position:absolute;inset:0;background:linear-gradient(90deg,rgba(0,0,0,.66) 0%,rgba(0,0,0,.38) 36%,rgba(0,0,0,.05) 70%,transparent 100%);pointer-events:none}.hero-copy{position:absolute;z-index:3;left:max(42px,calc((100vw - var(--max))/2 + 24px));top:50%;transform:translateY(-49%);width:min(700px,75vw);color:white}.hero-copy h1{font-family:var(--serif);font-size:clamp(3rem,5.5vw,5.2rem);line-height:1;margin:0 0 22px}.hero-copy .lead{font-size:clamp(1.35rem,2.25vw,2.05rem);line-height:1.25;font-weight:650;margin:0 0 28px}.keywords{font-weight:650;font-size:.95rem;letter-spacing:.015em}.hero-caption{position:absolute;z-index:3;right:max(30px,calc((100vw - var(--max))/2 + 24px));bottom:28px;color:white;text-align:right;font-size:.82rem;text-shadow:0 1px 4px #000}.dots{position:absolute;z-index:3;left:max(42px,calc((100vw - var(--max))/2 + 24px));bottom:30px;display:flex;gap:10px}.dot{width:8px;height:8px;border-radius:50%;background:rgba(255,255,255,.45);border:0;padding:0}.dot.active{background:#fff}
.section{padding:88px 0}.section-title,.page-title{font-family:var(--serif);font-weight:700}.section-title{font-size:clamp(2.2rem,3.5vw,3.6rem);margin:0 0 20px}.section-lead{font-size:1.14rem;color:var(--muted);max-width:1000px}.page-hero{padding:78px 0 46px}.page-title{font-size:clamp(3rem,5vw,5rem);line-height:1.05;margin:0 0 18px}.page-subtitle{font-size:1.45rem;color:var(--accent);font-weight:750;margin:0}.rule{border:0;border-top:1px solid var(--line);margin:0}
.research-cards{display:grid;grid-template-columns:repeat(4,1fr);gap:18px;margin-top:44px}.research-card{min-height:315px;border-radius:13px;padding:28px;display:flex;flex-direction:column}.research-card:nth-child(1){background:var(--card1)}.research-card:nth-child(2){background:var(--card2)}.research-card:nth-child(3){background:var(--card3)}.research-card:nth-child(4){background:var(--card4)}.research-card .num{font-size:.86rem;font-weight:800;color:white}.research-card h3{font-family:var(--serif);font-size:1.45rem;line-height:1.18;margin:30px 0 16px}.research-card p{font-size:.95rem;margin:0}.research-card .explore{margin-top:auto;padding-top:26px;font-size:.78rem;font-weight:800;letter-spacing:.06em}
.logo-strip{background:white;padding:54px 0}.logos{display:flex;align-items:center;justify-content:center;gap:100px;flex-wrap:wrap}.logos img{object-fit:contain;max-height:85px;max-width:220px}
.site-footer{background:#ebeae5;padding:42px 0;color:var(--muted);font-size:.9rem}.footer-flex{display:flex;justify-content:space-between;gap:30px;flex-wrap:wrap}.footer-brand{font-weight:800;color:var(--ink);letter-spacing:.05em}
.theme{padding:70px 0;border-top:1px solid var(--line)}.theme-grid{display:grid;grid-template-columns:minmax(0,.9fr) minmax(0,1.1fr);gap:58px;align-items:start}.theme-num{font-size:.8rem;font-weight:800;color:var(--accent)}.theme h2{font-family:var(--serif);font-size:2.45rem;line-height:1.15;margin:8px 0 18px}.theme-question{color:var(--accent);font-size:1.13rem;font-weight:750;margin-bottom:28px}.theme-copy p{font-size:1.03rem}.visual{background:var(--paper);border-radius:12px;padding:18px;box-shadow:0 10px 26px rgba(31,35,35,.05);overflow:hidden}.visual img,.visual video{width:100%;height:auto;border-radius:7px}.visual figure{margin:0;min-width:0}.visual figcaption{font-size:.76rem;color:var(--muted);line-height:1.35;margin-top:9px}.visual-grid-two{display:grid;grid-template-columns:1fr 1fr;gap:14px;align-items:center}.visual-grid-three{display:grid;grid-template-columns:1.05fr 1fr;gap:14px}.visual-stack{display:grid;gap:14px}.visual-caption{font-size:.78rem;color:var(--muted);margin-top:9px}
.theme01-grid{display:grid;grid-template-columns:.43fr .57fr;gap:18px;align-items:center}.theme01-grid figure:first-child img{max-height:405px;object-fit:contain}.theme01-grid figure:last-child img{max-height:470px;object-fit:contain}
.theme02-grid{display:grid;grid-template-columns:1.08fr .92fr;gap:16px;align-items:start}.theme02-right{display:grid;grid-template-rows:auto auto;gap:16px}.theme02-grid figure{background:#f2f0eb;border-radius:8px;padding:10px;overflow:hidden}.theme02-grid video,.theme02-grid img{display:block;width:100%;height:auto;object-fit:contain;border-radius:5px}.grooming-figure video{aspect-ratio:4/3;object-fit:cover}.theme02-right figure:first-child img{max-height:225px}.theme02-right figure:last-child img{max-height:270px}
.study-visual{background:#efede7}.study-visual figure{background:#fff;border-radius:8px;padding:10px}.theme03-grid{display:grid;grid-template-columns:1.22fr .78fr;gap:18px;align-items:stretch}.theme03-grid figure{display:grid;grid-template-rows:380px auto;align-items:start}.theme03-grid img{width:100%;height:380px;object-fit:contain}.theme04-grid{display:grid;grid-template-columns:1.08fr .92fr;gap:18px;align-items:center}.theme04-grid img{max-height:400px;object-fit:contain}
.button-link{display:inline-flex;align-items:center;gap:8px;margin-top:22px;padding:10px 15px;border:1px solid var(--accent);border-radius:999px;color:var(--accent);font-weight:800;font-size:.82rem}.button-link img{width:25px;height:25px;object-fit:contain}
.digital-section{padding:80px 0;background:var(--soft)}.digital-grid{display:grid;grid-template-columns:1fr 1fr;gap:26px;margin-top:36px;align-items:stretch}.digital-card{background:var(--paper);border-radius:14px;padding:26px;display:grid;grid-template-rows:auto auto auto 1fr;align-items:start}.digital-card h3{font-family:var(--serif);font-size:1.85rem;margin:0 0 6px}.digital-card .sub{color:var(--accent);font-weight:750;margin-bottom:20px}.figure-button{border:0;padding:0;background:white;cursor:zoom-in;width:100%;height:390px;display:flex;align-items:center;justify-content:center;border-radius:8px;overflow:hidden}.digital-card img{width:100%;height:100%;object-fit:contain;background:white}.digital-card p{font-size:.96rem;color:#3e4443;margin:22px 0 0}
.project-hero{padding:70px 0 48px}.project-top{display:grid;grid-template-columns:.92fr 1.08fr;gap:56px;align-items:start}.project-title{font-family:var(--serif);font-size:clamp(3rem,5vw,5rem);margin:0 0 18px}.project-question{font-size:1.55rem;color:var(--accent);font-weight:800}.project-logo{width:330px;max-height:100px;object-fit:contain;margin-left:auto;margin-bottom:18px}.concept img{width:100%;background:white;border-radius:10px}.concept-label{font-size:.76rem;color:var(--accent);font-weight:800;letter-spacing:.05em;margin-top:8px}.compare{display:grid;grid-template-columns:1fr 420px;gap:50px;align-items:start}.species{font-size:1.25rem;font-weight:800;color:var(--accent);word-spacing:6px}.compare img{width:100%;background:white;border-radius:10px;padding:8px}.team-strip{background:var(--paper);border-radius:12px;padding:22px 26px;margin-top:34px}.team-grid{display:grid;grid-template-columns:repeat(4,1fr);gap:18px}.team-name{font-weight:800}.team-affil{font-size:.8rem;color:var(--muted)}.team-collab{font-size:.83rem;color:var(--muted);margin:16px 0 0}.project-local{background:var(--soft);padding:72px 0}.project-cards{display:grid;grid-template-columns:repeat(3,1fr);gap:22px;margin-top:35px}.project-card{background:var(--paper);border-radius:13px;padding:24px}.project-card h3{font-family:var(--serif);font-size:1.5rem;margin:6px 0 14px}.project-card p{color:var(--muted);font-size:.93rem}.project-card img{margin-top:20px;width:100%;max-height:500px;object-fit:contain;background:white}.closing-question{font-family:var(--serif);font-size:2rem;line-height:1.35;margin-top:55px}
.people-feature{background:var(--paper);border-radius:14px;padding:28px;display:grid;grid-template-columns:390px 1fr;gap:56px;align-items:center}.people-feature img{width:100%;aspect-ratio:1/1;object-fit:cover;border-radius:10px}.people-feature h2{font-family:var(--serif);font-size:2.5rem;margin:0 0 10px}.role{color:var(--accent);font-weight:800}.affil{color:var(--muted)}.people-grid{display:grid;grid-template-columns:repeat(4,1fr);gap:26px}.person{background:var(--paper);border-radius:11px;padding:12px}.person img{width:100%;aspect-ratio:1.05/1;object-fit:cover;border-radius:7px}.person h3{font-size:1rem;margin:13px 3px 0}.person p{font-size:.8rem;color:var(--muted);margin:3px}.alumni-bg{background:var(--soft)}.alumni-grid{display:grid;grid-template-columns:repeat(5,1fr);gap:18px}.alumnus{background:var(--paper);border-radius:10px;padding:10px}.alumnus img{width:100%;aspect-ratio:1.1/1;object-fit:cover;border-radius:7px}.alumnus h3{font-size:.86rem;margin:10px 3px 2px}
.pub-year{display:grid;grid-template-columns:100px 1fr;gap:28px;margin-top:25px}.pub-year h2{font-family:var(--serif);font-size:1.8rem;color:var(--accent);margin:4px 0}.pub-list{border-top:1px solid var(--line)}.pub{display:grid;grid-template-columns:1fr 150px;gap:24px;padding:24px 0;border-bottom:1px solid var(--line)}.pub h3{font-size:1.05rem;line-height:1.4;margin:0 0 7px}.pub-authors{font-size:.78rem;color:var(--muted);margin:0}.pub-journal{font-family:var(--serif);font-style:italic;font-size:.84rem;margin:7px 0 0}.pub-link{text-align:right;color:var(--accent);font-weight:800;font-size:.75rem;padding-top:2px}.records{background:var(--paper);border-radius:12px;padding:26px 30px;margin-top:52px}.record-links{display:flex;gap:24px;margin-top:16px;color:var(--accent);font-weight:800;font-size:.82rem}
.about-copy p{font-size:1.05rem;max-width:1250px}.contact-card{background:var(--paper);border-radius:14px;padding:30px;display:grid;grid-template-columns:1.1fr .65fr 440px;gap:36px;align-items:start;margin-top:52px}.contact-card img{width:100%;max-height:260px;object-fit:cover;border:1px solid var(--line);border-radius:8px}.contact-links a{display:block;color:var(--accent);font-weight:800;font-size:.82rem;margin-bottom:14px}.email{color:var(--accent)}
.lightbox{position:fixed;inset:0;background:rgba(15,18,18,.92);z-index:100;display:none;align-items:center;justify-content:center;padding:34px}.lightbox.open{display:flex}.lightbox img{max-width:95vw;max-height:90vh;object-fit:contain;background:white}.lightbox-close{position:absolute;right:26px;top:18px;color:white;background:none;border:0;font-size:2rem;cursor:pointer}.sr-only{position:absolute;width:1px;height:1px;padding:0;margin:-1px;overflow:hidden;clip:rect(0,0,0,0);white-space:nowrap;border:0}
@media(max-width:1000px){.nav-links{display:none;position:absolute;left:0;right:0;top:77px;background:var(--paper);padding:22px 24px;flex-direction:column;align-items:flex-start;border-bottom:1px solid var(--line)}.nav-links.open{display:flex}.menu-btn{display:block}.research-cards,.project-cards{grid-template-columns:1fr 1fr}.theme-grid,.project-top,.compare{grid-template-columns:1fr}.compare{gap:24px}.digital-grid{grid-template-columns:1fr}.people-grid{grid-template-columns:repeat(3,1fr)}.alumni-grid{grid-template-columns:repeat(3,1fr)}.people-feature{grid-template-columns:300px 1fr}.contact-card{grid-template-columns:1fr 1fr}.contact-card img{grid-column:1/-1}.team-grid{grid-template-columns:1fr 1fr}.logos{gap:55px}}
@media(max-width:700px){.container,.narrow{width:min(calc(100% - 30px),var(--max))}.hero{min-height:640px}
.hero-slide:nth-child(2) img{position:absolute;right:1.5%;top:4%;width:42%;height:84%;object-fit:contain;object-position:center right}
.hero-slide:nth-child(3) img{position:absolute;left:auto;right:3%;top:1.5%;width:52%;height:87%;object-fit:contain;object-position:center right}
.hero-slide:nth-child(4) img{right:2%;top:31%;width:68%;height:46%}.hero-copy{left:24px;top:46%;width:calc(100% - 48px)}.hero-copy h1{font-size:3.2rem}.keywords{line-height:1.8}.hero-caption{right:20px;bottom:24px;max-width:60%}.dots{left:24px}.section{padding:62px 0}.research-cards,.project-cards,.people-grid,.alumni-grid,.team-grid{grid-template-columns:1fr}.theme-grid{gap:30px}.theme h2{font-size:2rem}.theme02-grid,.theme01-grid,.theme03-grid,.theme04-grid,.visual-grid-two,.visual-grid-three{grid-template-columns:1fr}.theme02-right{grid-template-rows:auto}.digital-card{grid-template-rows:auto auto 320px 1fr}.figure-button{height:320px}.project-logo{margin:0 0 18px}.people-feature{grid-template-columns:1fr}.people-feature img{max-width:360px}.pub-year{grid-template-columns:1fr;gap:2px}.pub{grid-template-columns:1fr}.pub-link{text-align:left}.contact-card{grid-template-columns:1fr}.logos img{max-width:170px}.record-links{flex-direction:column;gap:8px}.footer-flex{flex-direction:column}.page-hero{padding-top:55px}.page-title{font-size:3.2rem}}



/* In-page anchor offset for the sticky navigation. */
html{scroll-padding-top:74px}
.theme[id]{scroll-margin-top:74px}


/* Research figures use the existing lightbox; video remains a video rather than a still-image lightbox. */
.theme01-grid figure img,.theme02-right figure img,.theme03-grid figure img,.theme04-grid figure img{cursor:zoom-in}

.lightbox-content{display:flex;flex-direction:column;align-items:center;max-width:min(92vw,1400px);max-height:92vh}
.lightbox-content>img,.lightbox-video{max-width:92vw;max-height:82vh;object-fit:contain}
.lightbox-video{display:none;background:#000}
.lightbox-caption{margin-top:12px;color:#fff;font-size:14px;line-height:1.35;text-align:center;max-width:900px}
.lightbox-video-trigger{cursor:zoom-in}

/* OxytocINspace v0.11 refinements */
.project-hero-inner{position:relative;padding-top:74px}
.standalone-erc-logo{position:absolute;right:0;top:0;width:185px!important;height:auto!important;object-fit:contain;background:transparent!important;z-index:2}
.project-top{padding-top:0}
.team-heading{font-size:12px;font-weight:700;letter-spacing:.16em;color:var(--muted);margin:0 0 18px;text-transform:uppercase}
.project-grid .project-card:nth-child(2) img{display:block;width:calc(100% + 18px);max-width:none;margin-left:-18px;margin-right:0;margin-top:0;object-position:left top}
.project-grid .project-card img{vertical-align:top}


/* Align the Central Idea visual with the project question, not the main page title. */
.project-top>.concept{margin-top:102px}
@media(max-width:900px){.project-top>.concept{margin-top:24px}}

/* People page v0.14: bring the team into the initial viewport and add email affordances. */
.people-page-tight .page-hero{padding-bottom:24px}
.people-page-tight .section{padding-top:48px}
.people-page-tight .people-feature{grid-template-columns:320px 1fr;gap:44px;padding:22px}
.people-page-tight .people-feature .feature-portrait img{aspect-ratio:1/1}
.people-page-tight .people-feature h2{margin-top:0}
.people-page-tight .people-feature + *{margin-top:0}
.people-page-tight .people-feature-section{padding-top:16px;padding-bottom:36px}
.people-page-tight .collaborators-section{padding-top:32px}

.portrait-wrap{position:relative;overflow:hidden;border-radius:7px}
.feature-portrait{border-radius:10px}
.portrait-wrap img{display:block}
.person-mail{position:absolute;left:12px;bottom:12px;padding:8px 11px;border-radius:999px;background:rgba(18,24,24,.84);color:#fff;font-size:11px;font-weight:800;letter-spacing:.08em;text-decoration:none;opacity:0;transform:translateY(5px);transition:opacity .18s ease,transform .18s ease}
.portrait-wrap:hover .person-mail,.portrait-wrap:focus-within .person-mail{opacity:1;transform:translateY(0)}
.person-mail.pending{pointer-events:none}

.secondary-location{margin-top:22px;padding-top:18px;border-top:1px solid var(--line);max-width:620px}
.secondary-location p{margin:0 0 12px}
.secondary-location a{color:var(--accent);font-weight:700}

.digital-figure{margin:0;display:grid;grid-template-rows:390px auto}
.digital-figure .figure-button{height:390px}